Yıl: 2023 Cilt: 32 Sayı: 7 Sayfa Aralığı: 880 - 893 Metin Dili: Türkçe DOI: 10.55730/1300-0985.1880 İndeks Tarihi: 17-01-2024

More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye

Öz:
Fossil wood descriptions are among the most important indicators in understanding forest structures across geological ages. This study was carried out to obtain new findings about the Middle Eocene Climate Optimum (MECO) period of Anatolia with new samples collected from the Middle Eocene-aged fossil area near Amasya-Göynücek. The fossil area falls into the Göynücek Volcanics in the region. Transverse, radial, and tangential microscopic sections were taken from three new fossil wood samples, and all identifications were performed on these microscopic sections. As a result, the fossil species were identified as Pinuxylon cf. P. tarnocziense, cf. Dichrostachyoxylon zirkelii, and Laurinoxylon perseamimatus based on their wood anatomical features. In all three fossil woods, the growth rings are quite wide and their boundaries are indistinct. This indicates the existence of warm tropical conditions in the MECO period. On the other hand, the extant representatives of the angiosperm genera identified here are found in tropical-subtropical forests. Overall, more information was obtained about the forest structure and climate of tropical conditions in the MECO period of Anatolia.
Anahtar Kelime: Middle Eocene Climate Optimum Pinuxylon tarnocziense Dichrostachyoxylon zirkelii Laurinoxylon perseamimatus petrified wood Göynücek Volcanics

Belge Türü: Makale Makale Türü: Araştırma Makalesi Erişim Türü: Erişime Açık
  • Acarca Bayam NN, Akkemik Ü, Poole I, Akarsu F (2018). Further contributions to the early Miocene forest vegetation of the Galatean Volcanic Province, Turkey. Palaeobotanica Electronica 21: 21.3.40. https://doi.org/10.26879/816
  • Akgün F, Akay E, Erdoğan B (2002). Terrestrial to shallow marine deposition in Central Anatolia: a palynological approach. Turkish Journal of Earth Sciences 11: 1-27.
  • Akgün F, Kayseri MS, Akkiraz MS (2007). Palaeoclimatic evolution and vegetational changes during the Late Oligocene–Miocene period in Western and Central Anatolia (Turkey). Palaeogeography, Palaeoclimatology, Palaeoecology 253 (1-2): 56-90. https://doi. org/10.1016/j.palaeo.2007.03.034
  • Akgün F, Ocakoğlu F, Kayseri Özer MS, Michael W (2019). Preliminary palynological data from the Campanian Sakarya Archipelago: evidences for vegetation cover and paleo-environments. In: International Earth Science Colloquium on the Aegean Region; İzmir, Türkiye.
  • Akkemik Ü (2019). New fossil wood descriptions from Pliocene of central Anatolia and presence of Taxodioxylon in Turkey from Oligocene to Pliocene. Turkish Journal of Earth Sciences 28: 398-409. https://doi. org/10.3906/yer-1805-24
  • Akkemik Ü (2021). A re-examination of the angiosperm wood record from theearlyandmiddleMioceneofTurkey,andnewspeciesdescriptions. Acta Palaeobotanica 61 (1): 42-94. https://doi.org/10.35535/acpa- 2021-0004
  • Akkemik Ü, Akkılıç H, Güngör Y (2019). Fossil wood from the Neogene of the Kilyos coastal area in Istanbul, Turkey. Palaeontographica Abteilung B299 (1-6): 133-185. https://doi.org 10.1127/ palb/2019/0065
  • Akkemik Ü, Atıcı G, Poole I, Çobankaya M (2018). Three new silicified woods from a newly discovered earliest Miocene forest site in the Haymana Basin (Ankara, Turkey). Review of Palaeobotany and Palynology254:49-64.https://doi.org/10.1016/j.revpalbo.2018.04.012
  • Akkemik Ü, Güngör Y, Mantzouka D, Azaz D (2023). Mammeoxylon beylikduezuense Akkemik, Güngör, D. Mantzouka & Azaz sp. nov.: the first report of the genus for the Oligo/Miocene of Eurasia. Forestist 73(1): 28-41. https://doi.org/10.5152/forestist.2022.22024
  • Akkemik Ü, Iamandei S, Çelik H (2022). Further contribution to the early Miocene woody flora of Galatian Volcanic Province from Doğanyurt Village, Ankara (Turkey). Turkish Journal of Earth Sciences 31: 208- 234. https://doi.org/ 10.55730/1300-0985.1763
  • Akkemik Ü, Mantzouka D, Tunç U, Koçbulut F (2021). The first paleoxylotomical evidence from the Mid-Eocene Climate Optimum from Turkey. Review of Palaeobotany and Palynology 285. https:// doi.org/10.1016/j.revpalbo.2020.104356
  • Akkemik Ü, Sakınç M (2013). Sequoioxylon petrified woods from the Mid to Late Oligocene of Thrace (Turkey). IAWA Journal 34 (2): 177-182. https://doi.org/10.1163/22941932
  • Akkiraz MS, Akgün F, Örçen, S, Bruch A, Mosbrugger V (2006). Stratigraphic and palaeoenvironmental significance of Bartonian– Priabonian (Middle–Late Eocene) microfossils from the Başçeşme Formation, Denizli Province, Western Anatolia. Turkish Journal of Earth Sciences 15 (2): 155-180
  • Akkiraz MS, Sezgül Kayseri M, Akgün F (2008). Palaeoecology of coal-bearing Eocene sediments in Central Anatolia (Turkey) based on quantitative palynological data. Turkish Journal of Earth Sciences 17: 317-360.
  • Alessi N, Wellstein C, Spada F, Zerbe S (2018). Phytocoenological approach to the ecology of Laurus nobilis L. in Italy. Scienze Fisiche e Naturali, 29: 343-354. https://doi.org/10.1007/ s12210-021-00981-7
  • Biltekin D (2018). Palynomorphs from a lacustrine sequence provide evidence for palaeoenvironmental changes during the early Miocene in Central Anatolia, Turkey. Canadian Journal of Earth Sciences 55 (5): 505-513. https://doi.org/10.1139/cjes- 2017-0170
  • Boyer WD (1987). Volume growth loss: A hidden cost of periodic prescribed burning in longleaf pine? Southern Journal of Applied Forestry 11 (3): 154-157. https://doi.org/10.1093/ sjaf/11.3.154
  • Burke KD, Williams MA, Chandler AM, Haywood DJ, Lunt BL et al. (2018). Pliocene and Eocene provide best analogs for near-future climates. Proceedings of the National Academy of Sciences of the United States 115 (52): 13288-13293, https://doi. org/10.1073/pnas.180960011
  • Cleal CJ, Stolle E, van Waveren IM, King S, Didari V (2017). Carboniferous plant fossils from Northern Turkey in the Jongmans Collection, Naturalis, Leiden. Paleontological Journal 51 (7): 770-777. https://doi.org/10.1134/S0031030117070048
  • Denk T, Güner HT, Bouchal JM (2019). Early Miocene climate and biomes of Turkey: evidence from leaf fossils dispersed pollen, and petrified wood. Palaeogeography, Palaeoclimatology, Palaeoecology 530: 236-248. https://doi.org/10.1016/j. palaeo.2019.05.042
  • Dolezych M, Reinhardt L (2019). First evidence for the conifer Pinus, as Pinuxylon selmeierianum sp. nov., during the Paleogene on Wootton Peninsula, northern Ellesmere Island, Nunavut, Canada. Canadian Journal of Earth Science 57: 25-39. https:// doi.org/10.1139/cjes-2018-0163
  • Dupéron J, Dupéron-Laudoueneix M (1985). Considérations sur les gisements à végétaux tertiaires du Sud-Ouest de la France. Bulletin des Sciences Naturelles et de G éologie 8: 197-212 (in French).
  • Göçmengil G, Karacık Z, Genç ŞC, Billor MZ (2018). 40Ar-39Ar geochronology and petrogenesis of postcollisional trachytic volcanism along the İzmir-Ankara-Erzincan Suture Zone (NE, Turkey). Turkish Journal of Earth Sciences 27: 1-31. https:// doi.org/10.3906/yer-1708-4
  • Güner HT, Bouchal JM, Köse N, Göktaş F, Mayda S et al. (2017). Landscape heterogeneity in the Yatağan Basin (southwestern Turkey) during the middle Miocene inferred from plant macrofossils. Palaeontographica Abteilung B 296 (1-6): 113- 171. https://doi.org/10.1127/palb/2017/0057
  • Hassler M (2004–2023). World Plants. Synonymic Checklist and Distribution of the World Flora. Version 14.7; last updated on January 14, 2023. Available at www.worldplants.de.
  • Huard J (1966). Étude anatomique des bois de conifères des couches à lignite néogènes des Landes. Mémoires de la Société Géologique de France, Nouvelle Série 105: 1-85 (in French).
  • IAWA Committee (1989). IAWA list of microscopic features for hardwood identification. IAWA Bulletin 10: 219-232.
  • IAWA Committee (2004). IAWA list of microscopic features for softwood identification. IAWA Journal 25: 1-70.
  • InsideWood (2004–onwards). Published on the Internet. Available at http://insidewood.lib.ncsu.edu/search.
  • Jeong EK, Kim K, Suzuki M, Uemura K (2012). Daijima-type conifer wood assemblage of the Hatamura Formation (Middle Miocene) in the Akita Prefecture, Japan. Geosciences Journal 16 (2): 115-125. https://doi.org/10.1007/s12303-012-0018-3
  • Kayseri MS, Akgün F (2008).Palynostratigraphic, palaeovegetational and palaeoclimatic investigations on the Miocene deposits in Central Anatolia (Çorum Region and Sivas Basin). Turkish Journal of Earth Sciences 17 (2): 361-403.
  • Koçbulut F (2004). Neotectonic characteristics of the Ezinepazar- Sungurlu Fault in the northern part of Göynücek - Gediksaray Region (SW Amasya). PhD Thesis, Sivas Cumhuriyet University, Sivas, Türkiye (in Turkish with English abstract).
  • Koçbulut F, Kavak KŞ, Tatar O (2015). Analysis of Ezinepazarı- Sungurlu Fault Zone (Turkey) using Landsat TM data and its kinematic implications. Arabian Journal of Geosciences 8 (8): 6425-6439. https://doi.org/10.1007/s12517-014-1660-z
  • Kutluk H, Kır O, Akkemik Ü (2012). First report of Araucariaceae wood (Agathoxylon sp.) from the late Cretaceous of Turkey. IAWA Journal 33 (3): 316-326. https://doi.org/10.3195/ ejejfs.784543
  • Mantzouka D, Karakitsios V, Sakala J, Wheeler EA (2016). Using idioblasts to group Laurinoxylon species: case study from the Oligo-Miocene of Europe. IAWA Journal 37 (3): 459-488. https://doi.org/10.1163/22941932-20160147
  • Mantzouka D, Sakala J, Kvaček Z, Koskeridou E, Ioakim C (2019). Two fossil conifer species from the Neogene of Alonissos Island (Iliodroma, Greece). Geodiversitas 41 (3): 125-142. https://doi. org/10.5252/geodiversitas2019v41a3
  • Olson DM, Dinerstein E (1998). The Global 200: A representation approach to conserving the Earth’s most biologically valuable ecoregions. Conservation Biology 12: 502-515. https://doi. org/10.1046/j.1523-1739.1998.012003502.x
  • Özcan A, Erkan A, Keskin E, Oral A, Özer S et al. (1980). Kuzey Anadolu Fayı-Kırşehir Masifi Arasının Temel Jeolojisi: M.T.A. Derleme Rapor No: 6722. Ankara, Türkiye: MTA (in Turkish).
  • Petrescu J (1978). Etudes sur les flores paléogènes du nord-ouest de la Transsylvanie et de la Moldavie Central. Bucharest , Romania: University of Cluj-Napoca (in French).
  • Platzman ES, Platt JP, Tapirdamaz C, Sanver M, Rundle CC (1994). Why are there no clockwise rotations along the North Anatolian Fault Zone? Journal of Geophysical Research 99: 21705-21715. https://doi.org/10.1029/94JB01665
  • Privé C (1970). Sur un bois de Légumineuses du Stampien de Royat (Puy-de-Dôme). In: 94° Congrès national des sociétés savants; Pau, France. pp. 191-205 (in French).
  • Privé C (1972). Sur la présence de bois de pins fossiles dans le Cantal. In: Actes du 97 e congrès national des sociétés savantes. Nantes, France. pp. 137-158.
  • PROSEA (2023). Plant Resources of South East Asia. Available at https://prosea.prota4u.org/index.aspx.
  • Rössler W (1937). Pliozäne Koniferenhölzer der Umgebung von Gleichenberg in Steiermark. Mitteilungen des Naturwissenschaftlichen Vereines für Steiermark 74: 64-79 (in German).
  • Sayadi S (1973). Contribution à 1’étude de la flore miocène de laTurquie. Thesis, Paris, France (in French).
  • Selmeier A (1990). Dichrostachyoxlon zirkelii (Felix), Mimosoideae, a silicified wood from Miocene sediments of Küçük Çekmece Lake (Turkey). Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Histor Geologie 30: 121-135 (in German).
  • Tekeli O (1981). Subduction complex of Pre-Jurassic age, Northern Anatolia, Turkey. Geology 9: 68-72.
  • Van der Boon A, Kuiper FF, Van der Ploeg R, Cramwinckel MJ, Honarmand M (2021). Exploring a link between the Middle Eocene Climatic Optimum and Neotethys continental arc flare- up. Climate of the Past 7: 229-239. https://doi.org/10.5194/cp- 2020-48
  • Van der Burgh J (1964). Hölzer der niederrheinischen Braunkohlenformation, 1. Hölzer der Braunkohlengrube “Anna” zu Haanrade (Niederländisch Limburg). Acta Botanica Neerlandica 13 (2): 250-301 (in German).
  • Van der Burgh J (1973). Hölzer der niederrheinischen braunkohlenformation, 2. Hölzer der braunkohlengruben “Maria Theresia” zu herzogenrath, “Zukunft West” zu eschweiler und “Victor” (Zülpich mitte) zu Zülpich. Nebst einer systematischanatomischen bearbeitung der gattung Pinus L. Review of Palaeobotany and Palynology 15 (2): 73- 275 (in German).
  • Wang HB, Oskolski AA, Jacques FMB, Wang YH, Zhou ZK (2017). Lignified woods of Pinus (Pinaceae) from the late Miocene of central Yunnan, China, and their biogeographic and paleoclimatic implications. Palaeoworld 26: 553-563. https:// doi.org/10.1016/j.palwor.2016.06.003
  • Wheeler EA (2011). InsideWood - A web resource for hardwood anatomy. IAWA J. 32 (2), 199–211. https://doi. org/10.1163/22941932-90000051.
  • Yi TM (2002). Neogene fossil woods of Yunnan Province and their palaeoclimate implications. PhD, Institute of Botany, Chinese Academy of Sciences, Beijing, China (in Chinese with English abstract).
  • Yi TM, Li CS, Jiand XM (2005). Conifer woods of the Pliocene Age from Yunnan, China. Journal of Integrative Plant Biology 47 (3): 264-270. https://doi.org/10.1111/j.1744-7909.2005.00041.x
  • Zachos JC, Dickens GR, Zeebe RE (2008). An early Cenozoic perspective on greenhouse warming and carbon-cycle dynamics. Nature 451: 279-283. https://doi.org/10.1038/ nature06588
APA Akkemik Ü, Koçbulut F, Toprak Ö, TUNÇ U (2023). More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye. , 880 - 893. 10.55730/1300-0985.1880
Chicago Akkemik Ünal,Koçbulut Fikret,Toprak Özlem,TUNÇ UMUT More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye. (2023): 880 - 893. 10.55730/1300-0985.1880
MLA Akkemik Ünal,Koçbulut Fikret,Toprak Özlem,TUNÇ UMUT More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye. , 2023, ss.880 - 893. 10.55730/1300-0985.1880
AMA Akkemik Ü,Koçbulut F,Toprak Ö,TUNÇ U More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye. . 2023; 880 - 893. 10.55730/1300-0985.1880
Vancouver Akkemik Ü,Koçbulut F,Toprak Ö,TUNÇ U More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye. . 2023; 880 - 893. 10.55730/1300-0985.1880
IEEE Akkemik Ü,Koçbulut F,Toprak Ö,TUNÇ U "More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye." , ss.880 - 893, 2023. 10.55730/1300-0985.1880
ISNAD Akkemik, Ünal vd. "More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye". (2023), 880-893. https://doi.org/10.55730/1300-0985.1880
APA Akkemik Ü, Koçbulut F, Toprak Ö, TUNÇ U (2023). More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye. Turkish Journal of Earth Sciences, 32(7), 880 - 893. 10.55730/1300-0985.1880
Chicago Akkemik Ünal,Koçbulut Fikret,Toprak Özlem,TUNÇ UMUT More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye. Turkish Journal of Earth Sciences 32, no.7 (2023): 880 - 893. 10.55730/1300-0985.1880
MLA Akkemik Ünal,Koçbulut Fikret,Toprak Özlem,TUNÇ UMUT More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye. Turkish Journal of Earth Sciences, vol.32, no.7, 2023, ss.880 - 893. 10.55730/1300-0985.1880
AMA Akkemik Ü,Koçbulut F,Toprak Ö,TUNÇ U More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye. Turkish Journal of Earth Sciences. 2023; 32(7): 880 - 893. 10.55730/1300-0985.1880
Vancouver Akkemik Ü,Koçbulut F,Toprak Ö,TUNÇ U More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye. Turkish Journal of Earth Sciences. 2023; 32(7): 880 - 893. 10.55730/1300-0985.1880
IEEE Akkemik Ü,Koçbulut F,Toprak Ö,TUNÇ U "More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye." Turkish Journal of Earth Sciences, 32, ss.880 - 893, 2023. 10.55730/1300-0985.1880
ISNAD Akkemik, Ünal vd. "More evidence of tropical conditions in the Middle Eocene Climatic Optimum (MECO) with new fossil woods from North-Central Türkiye". Turkish Journal of Earth Sciences 32/7 (2023), 880-893. https://doi.org/10.55730/1300-0985.1880